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At the head of Parliament is the British Sovereign. The Sovereign's role, however, is merely ceremonial; in practice, he or she always acts on the advice of the Prime Minister and other ministers, who are in turn accountable to the two Houses of Parliament.
The Upper House, the House of Lords, is mostly made up of appointed members ("Lords of Parliament." The whole House is formally styled The Right Honourable The Lords Spiritual and Temporal in Parliament Assembled, the Lords Spiritual being clergymen of the Church of England and the Lords Temporal being Peers of the Realm. The Lords Spiritual and Lords Temporal are considered separate "estates," but they sit, debate and vote together.
The Lords Spiritual formerly included all of the senior clergymen of the Church of England—archbishops, bishops, abbots and priors. Upon the Dissolution of the Monasteries under Henry VIII, however, the abbots and priors lost their positions in Parliament. All diocesan bishops continued to sit in Parliament, but the Bishopric of Manchester Act 1847 and later acts provide that only the twenty-six most senior are Lords Spiritual. These twenty-six always include the incumbents of the "five great sees," namely, the Archbishop of Canterbury, the Archbishop of York, the Bishop of London, the Bishop of Durham and the Bishop of Winchester. The remaining twenty-one Lords Spiritual are the most senior diocesan bishops, ranked in order of consecration.
The Lords Temporal are all members of the Peerage. Formerly, they included hereditary peers, of the ranks of Duke, Marquess, Earl, Viscount and Baron. The right of some hereditary peers to sit in Parliament was not automatic: after Scotland and England united into Great Britain in 1707, it was provided that all peers whose dignities had been created by English Kings could sit in Parliament, but those whose dignities had been created by Scottish Kings were to elect a limited number of "representative peers." A similar arrangement was made in respect of Ireland when that nation merged with Great Britain in 1801. But when Southern Ireland left the United Kingdom in 1922, the election of Irish representative peers ceased. By the Peerage Act 1963, the election of Scottish representative peers also ended, but all Scottish peers were granted the right to sit in Parliament. Under the House of Lords Act 1999, only life peerage dignities (that is to say, peerage dignities which cannot be inherited) automatically entitle their holders to seats in the House of Lords. Of the hereditary peers, only ninety-two—the individuals exercising the offices of Earl Marshal and Lord Great Chamberlain, in addition to ninety hereditary peers elected by other peers—retain their seats in the House.
The Commons, the last of the "estates" of the Kingdom, are represented in the House of Commons, which is formally styled The Honourable The Commons in Parliament Assembled. The House consists of 659 members ("Members of Parliament" or "MPs"), each of whom is chosen by a single constituency according to the First-Past-the-Post electoral system. Universal adult suffrage prevails; citizens of the United Kingdom, as well as citizens of the Republic of Ireland and of Commonwealth nations resident in the United Kingdom, are qualified to vote. The term of members of the House of Commons depends on the term of Parliament; a general election, during which all the seats are contested, occurs after each dissolution (see below).
The three branches of Parliament are supposed to be kept separate from each other; no individual may form a part of more than one component of Parliament. Lords of Parliament are legally barred from voting in elections for members of the House of Commons; furthermore, the Sovereign by convention does not vote, although there is no statutory impediment.
Each of the two Houses of Parliament is presided over by a Speaker. In the House of Lords, the Lord Chancellor, a member of the Cabinet, is the ex officio Speaker. Where there is a vacancy in the office, a Speaker may be appointed by the Crown. Deputy Speakers, who take the place of an absent Lord Chancellor, are also chosen by the Crown.
The House of Commons has the right to elect its own Speaker. Theoretically, the approval of the Sovereign is required before the election becomes valid, but it is, by modern conventions, always granted. The Speaker's place may be taken by three deputies, known as the Chairman, First Deputy Chairman and Second Deputy Chairman of Ways and Means. (They take their name from the Committee of Ways and Means, of which they were once presiding officers, but which no longer exists.)
In general, the Lord Chancellor's influence as Speaker is very limited, whilst the powers belonging to the Speaker of the House of Commons are vast. Decisions on points of order and on the disciplining of unruly members are made by the whole body in the Upper House, but by the Speaker alone in the Lower House. Speeches in the House of Lords are addressed to the House as a whole (using the words "My Lords"), but those in the House of Commons are addressed to the Speaker alone (using the words "Mr Speaker" or "Madam Speaker").
Both Houses may decide questions with voice voting; members shout out "Aye" and "No" (in the House of Commons), or "Content" and "Not-Content" (in the House of Lords), and the presiding officer declares the result. The pronouncement of the Lord Chancellor or Speaker may be challenged, and a recorded vote (known as a division) demanded. (The Speaker of the House of Commons may choose to overrule a frivolous request for a division, but the Lord Chancellor does not possess an equivalent power.) In each House, a division requires members to file into one of the two lobbies alongside the Chamber; their names are recorded by clerks, and their votes are counted as they exit the lobbies to re-enter the Chamber. The Speaker of the House of Commons, who is expected to remain non-partisan, does not cast a vote except in the case of a tie; the Lord Chancellor, however, votes along with the other Lords.
(For further details on procedure, see the separate articles on the House of Lords and the House of Commons.)