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Leeches

Scientific classification
Kingdom:Animalia
Phylum:Annelida
Class:Clitellata *
Subclass:Hirudinea
Orders

Arhynchobdellida
Rhynchobdellida

*There is some dispute as to
whether Hirudinea should be a class
itself, or a subclass of the Clitellata.

A leech is an annelid in the subclass Hirudinea. There are freshwater, terrestrial and marine leeches. Like their near relatives, the Oligochaeta, they share the presence of a clitellum . Many species of leech are haemophagic parasites, living on occasional meals of blood obtained by attaching themselves to fish, amphibians (frogs etc.), and mammals. The medicinal leech, Hirudo medicinalis, which is native to Europe, has been used for clinical bloodletting for thousands of years. Most leech species are predators and feed on small worms or other invertebrates.

The leeches are clitellate when breeding and are derived from some oligochaete ancestor. They attach to their hosts and remain there. Leeches' bodies are composed of 32 segments. They all have a ventral sucker formed from the last six segments of the body, and that is a combination of mucus and suction caused by concentric muscles formed from the circular muscles of the segments from which the sucker is derived. Like earthworms, leeches are hermaphrodites.

Some species of leech will nurture their young, providing food, transport, and protection, which is unusual behavior in an invertebrate.[1]

1 Diversity

All leeches are from one of three orders:

  1. Rhynchobdellae (comprising leeches with an eversible proboscis)
    The Rhychobdellae consist of two families: The Glossiphoniidae (flattened leeches with a poorly developed anterior sucker) and the Pisciolidae (have cylindrical bodies and usually well-marked anterior sucker). The Glossiphoniidae are in fresh-water habitat, the Pisciolidae are found in both fresh-water and sea-water habitats.
  2. Gnathobdellae (leeches like the medicinal leech and horse leech which have cutting pharyngeal teeth)
    The Gnathobdellae has two main families. The Hirudidae includes the medicinal and horse leeches. The other family is Haemadipsidae, an tropical and subtropical group which hang themselves in wet forests and attach to passing animals.
  3. Pharyngobdellae (leeches have a sucking pharynx like that of ganthobdelliforms, but from which the teeth are absent)
    The Pharyngobdellae have six to eight pairs of eyes, as compared with five paris in gnathobdelliform leeches, and include three realated families. The Erpobdellidae are some species from freshwater habitats.

2 Use of Hirudo medicinalis in medicine

The leech has long been used in medicine, although today its use is mainly limited to limb reattachment procedures instead of the wide-ranging medical use of the past. Leeches were once so commonly used that doctors were popularly called leeches. In Old High German, lāhhi ( etymon of leech) means "physician".

Leech saliva contains a number of compounds which assist in its feeding. An anaesthetic limits the sensations felt by the host (and thus reduces the chance of the host trying to detach the leech). A vasodilator causes the blood vessels near the leech to become dilated, and thus provide the leech with a better supply.

Lastly, the leech saliva contains a complex protein called hirudin , which is a highly effective anticoagulant. The leech needs this to prevent blood clots (which would block its feeding) from forming in the wound created by its mouthparts. These properties are difficult to achieve using other medical techniques, and it is for this reason that leeches have come back into clinical practise in the last 25 years. The small amounts of hirundin present in leeches makes it unsuitable to be harvested for more general medical use, so hirudin (or related chemicals) have been synthesised using recombinant-DNA technology.

Bdellatomy is the practice of cutting the leech open slightly while it is sucking blood to let the blood in it out, so thinking that it is not full yet, the leech continues to bite instead of detach itself. This practise was first recorded in 1868 by Daily News .

3 The anatomy of medicinal leeches

The anatomy of medicinal leeches may look simple, but more details are found beyond the macro level. Externally, medicinal leeches tend to have a brown and red stripped design on an olive colored background. These organisms have two suckers, one at each end, called the anterior and posterior sucker. The posterior is mainly used for leverage while the anterior sucker, consisting of the jaw and teeth, is where the feeding take place. Medicinal leeches have three jaws that look like little saws, and on them are about 100 horny teeth used to incise the host.





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