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Plant improvement has been practiced for thousands of years. Domestication, classical plant breeding and genetic engineering are all processes that alter the genome of a plant to enhance its qualities as a crop.

Desirable traits for crop species include:

  1. Increased quality and yield of the crop
  2. Increased tolerance of environmental pressures ( salinity, extreme temperature, drought)
  3. Resistance to viruses, fungi and bacteria
  4. Increased tolerance to insect pests
  5. Increased tolerance of herbicides

Plant improvement is practiced worldwide and is extremely improtant for ensuring food security and developing practices of sustainable agriculture

1 Domestication

Domestication of plants is selectionIn the context of evolution, certain traits or alleles of a species may be subject to selection. Under selection, individuals with advantageous or "adaptive" traits tend to be more successful than their peers reproductively--meaning they contribute more o process conducted by humans to produce plants that meet the needs of the farmer and the consumer. Domestication of plants has been carried out for 9000 - 11 000 years, many crops in present day cultivation are the result of domestication in ancient times, about 5 000 years ago in the Old WorldThe Old World consists of those parts of Earth known to Europeans before the voyages of Christopher Columbus: Europe, Asia, and Africa. The term is in distinction for the New World, meaning the Americas. Although the interiors of Asia and Africa were not and 3000 years ago in the New WorldThe New World is one of the names used for the continents of North and South America and adjacent islands collectively, in use since the 16th century. The continents were new to the Europeans, who knew the world consisting only of Europe, Asia, and Africa, essentially all our important food crops had been domesticated. In the NeolithicThe Neolithic (Greek neos new, lithos stone, or "New Stone Age") is traditionally the last part of the stone age. The name was invented by John Lubbock in 1865 as a refinement of the three-age system. It followed Pleistocene Epipalaeolithic and early Holo period domestication took a minimum of 1000 years and a maximum of 7000 years.

2 Classical plant breeding

Classical plant breeding uses interbreeding (crossing) of closely or distantly related species to produce new cops with desirable properties. Plants are crossed to introduce traits/genes from one species into a new genetic background. For example a mildew resistant pea may be crossed with a high-yielding but susceptible pea, the goal of the cross would be to introduce the mildew resistance without losing the high-yield characteristics. Classical breeding relies on homologous recombinationGenetic recombination is a general category for a number of processes involving DNA studied in classical genetics, population genetics, molecular biology, and evolutionary biology. Crossing over Main article Chromosomal crossover The most important and we of two genomes to generate genetic diversity. It also makes use of a number of molecular techniques to generate diversity and produce plants that would not exist in nature.

The Yecoro wheataestivum ''T. aethiopicum ''T. araraticum ''T. boeoticum ''T. carthlicum ''T. compactum ''T. dicoccon ''T. durum ''T. ispahanicum ''T. karamyschevii ''T. militinae T. monococcum ''T. polonicum T. spelta ''T. timopheevii ''T. trunciale ''T. turanicum ''T. (right) cultivar is sensitive to salinity, plants resulting from a hybrid cross with cultivar W4910 (left) show greater tolerance to high salinity

When distantly related species are crossed plant breeders make use of a number of plant tissue culture techniques to produce progeny from other wise fruitless mating. Interspecific and intergeneric hybrids are produced from a cross of related species or genera that do not normally sexually reproduce with each other. The cereal triticale, is a wheat and rye hybrid. The first generation created from the cross was sterile, so the cell division inhibitor colchicine was used to double the number of chromosomes in the cell. Cells with an uneven number of chromosomes are sterile.

Failure to produce a hybrid may be due to pre- or post fertilization incompatibility. If fertilization is possible between two species or genera, the hybrid embryo aborts before maturation. When the cross is incompatible after fertilization the resulting embryo resulting from an interspecific or intergeneric cross can be rescued and cultured to produce a whole plant. This technique has been used to produce new rice for Africa, an interspecific cross of Asian rice (Otyza sativa) and African rice (Otyza glaberrima).

Chemical mutagens like EMS and DMSO, radiation and transposons are used to generate mutants with desirable traits to be bred with other cultivars. Classical plant breeders also generate genetic diversity within a species by exploiting a process called somaclonal variation. Somaclonal variation occurs in plants produced from tissue culture, particularly plants derived from callus.

When a desirable trait has been bred into a species, a number of crosses to the favoured parent are made to make the new plant as similar as the parent as possible. Returning to the example of the mildew resistant pea being crossed with a high-yielding but susceptible pea, to make the mildew resistant progeny of the cross most like the high-yielding parent, the progeny will be crossed back to that parent for several generations. This process removes most of the genetic contribution of the mildew resistant parent. Classical breeding is therefore a cyclical process.

It should be noted that the breeder does not know exactly what genes have been introduced to the new cultivars, and some scientists argue that plants produced by classical breeding methods should undergo the same safety testing regime as genetically modified plants.





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